Legendary Researchers Transform 
Science and Medicine 
T H E N : 
Our 2006 issue featured CU Boulder’s Marvin 
Caruthers, Ph.D., and University of Colorado geneticist 
Theodore Puck, Ph.D., whose discoveries advanced 
human genetics, cancer research, and biotechnology.
N OW: 
Caruthers’ DNA synthesis chemistry helped transform medicine and 
underpins modern genomics. He cofounded Amgen and Applied 
Biosystems, both launched from research developed at CU Boulder. Puck’s 
legacy includes major contributions to Colorado’s leadership in oncology 
research and therapies. His pioneering methods for growing colonies of 
human cells enabled early tests for radiation damage and later research into 
the genetic causes of cancer.
THEN & [NOW]
Celebrating 20+ Years of BioScience Colorado Magazine!
Visitors see how Medtronic’s Colorado 
team advances technologies used by 
clinicians and patients around the world.
2026-27 BIOSCIENCE COLORADO     11
Transforming Patient Care and Outcomes  
Through Personalized Diagnostics
A proud member of the Colorado Bioscience Community for 20+ years.  
Louisville, CO is home to our Global HQ and one of our CLEP | CAP accredited Labs.
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Supporting clinical decisions to expedite 
personalized care and improve  
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Enabling the world’s leading 
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